Age:
Westphalian D to Lower Permian
Classification:
FILICOPSIDA PICHI-SERMOLLI, MARATTIALES ENGLER & PRANTL.
Data
base: In total 1,100 sterile and 500 fertile pecopterids from Sydney
Basin, Nova Scotia Canada - Plzeň, Kladno-Rakovnik, Krkonoše
Piemont, and Intrasudetic Basins, Czech Republic.
Preservation:
Compression/impression, natural maceration, moderate maturation level
(vitrinite reflectance Ro% 0.7%, reaching locally 1%). Cuticle is studied
for the first time: recovery ratio better than 60%.
Unrevised
Canadian pecopterid species:
P. abbreviata P. acadica P. arborescens P. candolleana P. clarkii
P. cyathea
P. herdii P. lamuriana P. micromiltonii P. miltonii P. paleacea P. pectinata
P.
polymorpha P. robbii P. sterzeliformis P. untia. (P.
plumosa,
nonmarattialean).
Unrevised
Czech pecopterid species:
Asterotheca
alpina Acitheca ambigua Ast. arborescens Ast. aspidioides
Ast.
candolleana Ast. cyathea Ast. densifolia Ast. hemitilioides Ast. imbricata
Ast.
lepidorhachis Ast. miltonii Ast. nyranensis Ast. permica Ast. potoniei
Aci.
polymorpha Ast. raconensis Ast. thuringiaca Ptychocarpus unitus.
We have
larger fertile higher-order rachides specimens consisting of at least
2-order pinnae (ultimate and penultimate pinnae). Pinnules up to 10
mm long, 2-3 mm wide, attached to a rachis by entire base, or constricted;
margins entire, straight, parallel, or slightly convex; midveins well
developed; laterals veins simple or forked, straight or curved; ultimate
and higher-order rachides striate, punctate (trichome bases), scaly,
or with well-preserved tracheids.
Conclusion:
Species are completely separable based on stomatal type, e.g. :
P. arborescens (paracytic), P. aspidioides (cyclocytic), P. polypodioides
(anomocytic). Most common is P. polymorpha (monocyclocytic).
In-situ
spore recovery: 70%
Comparison:
To known sporae dispersae
Groups: Laevigatosporites-Punctatosporites 11-38 μm
Punctatosporites-Torispora
28-51 μm
Punctatisporites
34-76 μm
Cyclogranisporites
33-110 μm
Most complete studied group to date: P. polymorpha
Range:
34(82)116 μm
Species: Verrucosisporites microtuberosus, Punctatisporites
sp.
Cyclogranisporites sp.
Conclusion: Most conservative spore is from P. arborescens,
most diverse from P. Polymorpha. It is not possible to determine pecopterid
species from in situ spores. Distribution of fertile pinnules
is similar to living ferns.